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All classes of steroid hormones are synthesized from the common precursor cholesterol. The synthesis of sex steroid hormones is carried out under the catalysis of a series of enzymes, and the genes encoding these enzymes include hsd3b1 , hsd11b2 , hsd17b1 , hsd17b3 , and hsd20b2 of the beta-hydroxysteroid dehydrogenases (β-HSDs) superfamily, and cyp11a1 , cyp11b , cyp17a1 , cyp19a1a , and cyp21a1 of the cytochrome P450 (cP450) superfamily (Tokarz et al., 2015). Hsd3b1 is responsible for the oxidation and isomerization of Δ5-3β-hydroxysteroid precursors in order to form Δ4-3-ketosteroid (Nakamoto et al., 2012). Hsd11b2 catalyzes the conversion of cortisol to cortisone (Fuller et al., 2012), and also plays a crucial role in the conversion of 11β-OH-androstenedione to 11-ketotestosterone (11-KT) (Tokarz et al., 2015), which affects the production of the most important androgen in teleost fish. Furthermore Hsd17b1 converts inactive estrone to active estradiol (Mindnich et al., 2004), Hsd17b3 is an essential enzyme for the synthesis of 11-KT (Mindnich et al., 2005) and Hsd20b2 converts 17alpha-hydroxyprogesterone to 17alpha,20beta-DP, which has been identified as the maturation-inducing hormone of several teleosts (Senthilkumaran et al., 2002). Materials and Methods. Genome-Wide Identification of β -hsd Genes. Phylogenetic Analysis. Gene Expression Profiling by Quantitative Real-Time PCR (qRT-PCR) A total of 22 and 19 β -hsd genes were collected from the human and zebrafish genome, respectively, which were used to construct HMM models. A set of 19 putative members of the β -hsd genes have been identified from the orange-spotted grouper genome (Table 1). Among the 19 β -hsd genes, a total of 14 were distributed across 10 linkage groups (LGs) in the grouper genome (Figure 1). In addition, we also identified 24, 16, 24, 17, 14, 19, 19, 19, 17, and 27 β -hsd genes in mouse, chicken, xenopus, coelacanth, spotted gar, fugu, tilapia, medaka, stickleback, and common carp, respectively (Table 1). The gene and protein IDs of the β -hsd genes of all species are listed in the attached table (Supplementary Table S1).
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